MALACOLOGIA
http://www.instituteofmalacology.org
One of the World’s Leading Malacological Journals
Journal Impact Factor 13.5 (2019)
Founded in 1961
OFFICERS
Editor-in-Chief
REBECCA J. RUNDELL
Associate Professor
Head Curator
Roosevelt Wild Life Collections
SUNY-ESF | State University of New York College of Environmental Science and Forestry
Syracuse, New York USA
https://www.snailevolution.org/
Malacologia E-mail Address: rundellmollusca@gmail.com
Editorial Office
CHARLES LYDEARD
Morehead State University
Department of Biology and Chemistry
103 Lappin Hall
Morehead, Kentucky 40351, U.S.A.
clydeardmalac@gmail.com
Composition Editor
PAULA M. MIKKELSEN
Field Museum of Natural History
Chicago, Illinois
Honorary Associate Editor
GEORGE M. DAVIS
Business Manager
DAVID CAMPBELL
110 S Main St, Box 7236
Boiling Springs, North Carolina 28017, U.S.A
malacologia@gardner-webb.edu
Copy Editor
EUGENE COAN
Santa Barbara Museum of Natural History
Santa Barbara, California, U.S.A.
genecoan@gmail.com
Advisory Editor
PAULA M. MIKKELSEN
THOMAS WILKE
Justus Liebig University
Giessen, Germany
tom.wilke@allzool.bio.uni-giessen.de
MALACOLOGIA is published by the INSTITUTE OF MALACOLOGY, U.S.A.
BOARD OF DIRECTORS
President
ELLEN STRONG
National Museum of Natural History
Washington D.C.
Vice President
RÜDIGER BIELER
Field Museum of Natural History, Chicago, Illinois
Treasurer
CHARLES LYDEARD
Morehead State University
Morehead, Kentucky
Secretary
PAULA M. MIKKELSEN
Field Museum of Natural History, Chicago, Illinois
Members
RÜDIGER BIELER
Field Museum of Natural History Chicago, Illinois
CAROLE S. HICKMAN
University of California, Berkeley, California
DIARMAID Ó FOIGHIL
University of Michigan, Ann Arbor,
Michigan
Copyright © 2023 by the Institute of Malacology
ISSN: 0076-2997 (print)
ISSN: 2168-9075 (Online)